Well, in this case its MKV. Also notice the double subtitle. Totem would tell you about a plugin available to download to play the content. A rather nice and useful feature IMO. I think in latest Ubuntu version you even can buy Fluendo codec packs directly from Ubuntu.
They're maemo and MeeGo... "Meamo!" sounds like what Zorro would say to catherine zeta jones... after she slaps him for looking at her dirtily...